Cut-paper reconstruction of a small four-legged early relative of mammals in side view, low and long-bodied, with short legs and a stubby tail

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The trackmaker

Plants and animalsearly relative of mammals (reconstruction)

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What it is

The animal that left the footprints at Corncockle Muir, in Dumfriesshire, Scotland, the first fossil footprints ever described by science. Nobody has found its bones, so the chapter shows a reconstruction. The tracks themselves are named Chelichnus.

Climbing a dune

The sandstone formed from desert dunes in the Permian. The quarrymen told Henry Duncan the tracks always ran up or down the sloping beds. On the way up, the claws dug in deeper, and each step pushed a small crescent of sand out behind the heel. Each front print shows up to five short toes on one rounded sole, about two inches across.

Not a tortoise

William Buckland matched the prints to his tortoises. A modern 3D scan of the Chelichnus duncani type slab found a ridge between the tip and the base of the toe marks. That is one reason experts now class these tracks with the therapsids, early relatives of mammals.

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